1. Department of Zoology, University of Oxford, Oxford OX1 3PS, UK
2. Institute of Integrative Biology, ETH Zürich, 8092, Zürich, Switzerland
3. Department of Biology, 208 Mueller Laboratory, Pennsylvania State University, University Park, PA 16802, USA
4. Department of Biosciences, University of Exeter, Cornwall Campus, Penryn TR10 9EZ, UK
5. Commonwealth Scientific and Industrial Research Organization Ecosystem Sciences and Sustainable Agriculture Flagship, Box 284, Canberra, Australian Capital Territory 2601, Australia