1. Department of Biology, University Brunei Darussalam, Tungku Link, Gadong 1410, Brunei Darussalam
2. Department of Animal Ecology and Tropical Biology, University of Würzburg, Biozentrum, Am Hubland, 97074 Würzburg, Germany
3. Zoological Institute and Museum, Greifswald University, Johann-Sebastian-Bach-Strasse 11/12, 17489 Greifswald, Germany