1. Senckenberg Natural History Collections Dresden, Museum of Zoology, Dresden, Germany
2. Division of Fishes, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA
3. Naturhistorisches Museum Bern, 3005 Bern, Switzerland
4. Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland