1. College of Life and Environmental Sciences, University of Exeter, Exeter, UK
2. Environment and Sustainability Institute, University of Exeter, Penryn Campus, Penryn, Cornwall, UK
3. Department of Zoology, University of Oxford, Oxford, UK
4. School of Biological Sciences, University of Bristol, Bristol, UK
5. Department of Genetics, Evolution, and Environment, University College London, London, UK
6. Departmento de Biogeografía y Cambio Global, Museo Nacional de Ciencias Naturales, Madrid E28006, Spain