1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, People's Republic of China
2. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China
3. Unidad Ejecutora Lillo, Conicet-Fundación Miguel Lillo, Miguel Lillo 251, Tucumán, Argentina
4. Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 2050, South Africa