1. Department of Aquatic Resources, Institute of Freshwater Research, Swedish University of Agricultural Sciences, Stångholmsvägen 2, 178 93 Drottningholm, Sweden
2. Department of Aquatic Resources, Swedish University of Agricultural Sciences, Box 7018, 750 07 Uppsala, Sweden
3. Biodiversity Unit, University of Turku, 20014 Turku, Finland
4. Department of Biology, University of Turku, 20014 Turku, Finland