1. Department of Biology, Western University , London, Ontario N6A 5B7, Canada
2. Department of Biology, Memorial University of Newfoundland , St John’s, Newfoundland A1C 5S7, Canada
3. Department of Biological Sciences, University of Cincinnati , Cincinnati, OH 45221, USA
4. Department of Biological Sciences, University of Alberta , Edmonton, Alberta T6G 2E9, Canada