1. Southwest Fisheries Science Center, National Marine Fisheries Service, NOAA , La Jolla, CA 92037, USA
2. University of British Columbia , Vancouver, British Columbia V6T 1Z4, Canada
3. Marine Mammal Institute, Oregon State University , Newport, OR 97365, USA
4. Northwest Fisheries Science Center, National Marine Fisheries Service, NOAA , Seattle, WA 98112, USA