1. Centre for Ecology, Evolution, and Conservation, School of Biological Sciences, University of East Anglia, Norwich NR4 7TJ, UK
2. Population Genetics Group, Institute of Marine Research, 5817 Bergen, Norway
3. Faculty of Biological Sciences, School of Biology, University of Leeds, Leeds LS2 9JT, UK
4. Centre for Environment Fisheries and Aquaculture Science (CEFAS), Lowestoft NR33 0HT, UK