1. Conservation Biology Group, Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
2. Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, UK
3. Department of Ecology and Evolutionary Biology, Eno Hall, Princeton University, Princeton, NJ 08544-1003, USA
4. Royal Society for the Protection of Birds, The Lodge, Sandy, Bedfordshire SG19 2DL, UK
5. Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, UK