1. NERC Centre for Population Biology, Division of Biology, Imperial College LondonSilwood Park, Ascot, Berkshire SL5 7PY, UK
2. Département de Biologie, Faculté des Sciences, Université de SherbrookeSherbrooke, Quebec J1K 2R1, Canada
3. Redpath Museum and Department of Biology, McGill University859 Sherbrooke Street West, Montreal, Quebec H3A 2K6, Canada