1. 1Department of Neurology, Massachusetts General Hospital, Fruit Street, Boston, MA 02114, USA
2. Medical and Molecular Genetics, GKTMedical and Dental School, Kings College, 8th Floor, Guy'sTower, London SE1 9RT, UK
3. Department of Anatomy and Developmental Biology, University College London, Gower Street, LondonWC1E 6BT, UK