1. Organic Geochemistry Unit, Bristol Biogeochemistry Research Centre, and The Cabot Institute, School of Chemistry, University of Bristol, Cantock's Close, Bristol BS8 1TS, UK
2. School of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, Bristol BS8 1RJ, UK
3. Alfred Wegener Institute for Polar and Marine Research, Am Alten Hafen 26, Bremerhaven 27568, Germany