1. Department of Biology, McGill University, Montreal, Canada H3A 1B1
2. Quebec Centre for Biodiversity Science, Montreal, Canada H3A 1B1
3. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig 04103, Germany
4. Institute of Computer Science, Martin Luther University Halle-Wittenberg, Halle (Saale) 06099, Germany
5. Department of Zoology and Biodiversity Research Centre, University of British Columbia, Vancouver V6T 1Z4, Canada
6. Santa Fe Institute, Santa Fe, NM 87501, USA