1. School of Geography and Tourism, Qufu Normal University, Rizhao 276826, People's Republic of China
2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Centre for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, Nanjing 210008, People's Republic of China
3. University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China