1. Centre for the Advanced Study of Collective Behaviour, University of Konstanz, Konstanz, Baden-Württemberg, 78464, Germany
2. Ecology of Animal Societies Department, Max Planck Institute of Animal Behavior, Radolfzell, Baden-Württemberg, 78315, Germany
3. Collective Behavior Department, Max Planck Institute of Animal Behavior, Radolfzell, Baden-Württemberg, 78315, Germany
4. Integrative Biology Department, Michigan State University, East Lansing, Michigan, 48824, USA