1. Department of Natural Sciences, National Museums Scotland, Edinburgh EH1 1JF, UK
2. Institute of Geography, School of Geosciences, University of Edinburgh, Edinburgh EH8 9YL, UK
3. Institute of Tropical Biodiversity and Sustainable Development, University Malaysia Terengganu, Kuala Nerus, Terengganu 21030, Malaysia
4. Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, The Recanti-Kaplan Centre, Tubney House, Abingdon Road, Abingdon, Oxfordshire OX13 5QL, UK
5. Zoological Museum, M.V. Lomonosov Moscow State University, Bolshaya Nikitskaya 2, Moscow 119991, Russia
6. Federal Scientific Centre for the Biodiversity of Terrestrial Biota of East Asia, Far Eastern Branch, Russian Academy of Sciences, Vladivostok, Primorskij kraj, Russia
7. Human Ecodynamics Research Centerand Doctoral Program in Anthropology, City University of New York (CUNY), NY 10017, USA