1. Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA
2. Natural History Museum of Geneva, Department of Mammalogy and Ornithology, CP 6434, CH-1211 Geneva 6, 6434, Switzerland
3. Muséum National d'Histoire Naturelle, Département Systématique et Evolution, UMR7205, Case Postale 51, 55 Rue Buffon, 75005 Paris, France
4. American Museum of Natural History, Center for Biodiversity and Conservation, Central Park West at 79th Street, New York, NY 10024, USA