1. College of Fisheries and Ocean Sciences, University of Alaska Fairbanks, Fairbanks, AK 99775, USA
2. South African Institute for Aquatic Biodiversity, Makhanda 6140, South Africa
3. Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520, USA
4. College of Science and Engineering, James Cook University, Townsville, Queensland 4811, Australia
5. Biodiversity and Geosciences Program, Museum of Tropical Queensland, Queensland Museum, Townsville, Queensland 4810, Australia
6. Department of Biological Sciences and Museum of Natural Science, Louisiana State University, Baton Rouge, LA 70803, USA
7. Yale Peabody Museum of Natural History, Division of Vertebrate Zoology. New Haven, CT 06520, USA