1. Department of Entomology, Penn State University University Park, PA, USA
2. Clemson University Arthropod Collection Department of Plant and Environmental Sciences 277 Poole Agricultural Center Clemson University, Clemson, SC 29634-0310, USA
3. Negaunee Integrative Research Center, Field Museum of Natural History 1400 S. DuSable Lake Shore Drive Chicago, IL 60605, USA