1. Australian National Herbarium, CPBR, CSIRO Plant Industry, GPO Box 1600, Canberra, ACT, 2601, Australia; Division of Botany and Zoology, Australian National University Canberra, ACT 2601 Australia
2. Australian National Herbarium, CPBR, CSIRO Plant Industry, GPO Box 1600 Canberra, ACT 2601 Australia
3. Division of Botany and Zoology, Australian National University Canberra, ACT 2601 Australia
4. School of Tropical Biology, James Cook University, Cairns, Queensland 4870 Australia