1. Departamento de Botánica; Universidad de Concepción; Casilla 160-C, Concepción Chile
2. Department of Ecology and Evolutionary Biology; University of Kansas; Lawrence, Kansas 66047 U.S.A.
3. Department of Higher Plant Systematics and Evolution; Institute of Botany, University of Vienna; Rennweg 14, A-1030, Vienna Austria
4. Departamento de Biología Molecular; Universidad de Concepción; Casilla 160-C, Concepción Chile