1. 1 Anses, INRA, ENVA, UPEC, UMR 1161 Virology, 23 avenue du général du Gaulle, 94706 Maisons-Alfort, France
2. 2 IRVT, Tunis-La Rabta, Tunisia
3. 4 Faculty of Sciences of Bizerte, Jarzouna Bizerte, Tunisia — under the tutelage of Carthage University, Tunisia
4. 3 ENMV, Sidi Thabet, Tunis, Tunisia — under the tutelage of Manouba University, Tunisia