1. Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, CO 80523, USA
2. Present address: Department of Statistics, University of Missouri, Columbia, MO 65211, USA
3. U.S. Department of Agriculture-Wildlife Services-National Wildlife Research Center, 4101 LaPorte Avenue, Fort Collins, CO 80521, USA
4. The Nature Conservancy, Global Lands Team, 117 E Mountain Avenue, Suite 201, Fort Collins, CO 80524, USA
5. Colorado Parks and Wildlife, 0088 Wildlife Way, Glenwood Springs, CO 81601, USA