1. School of the Environment, Washington State University , 1230 Webster Hall, Pullman, Washington 99164 , USA
2. Department of Forestry and Natural Resources, University of Kentucky , 730 Rose Street, Lexington, Kentucky 40546 , USA
3. Wildlife Health Program, Kentucky Department of Fish and Wildlife Resources , 1 Sportsman’s Lane, Frankfort, Kentucky 40601 , USA
4. Bear and Elk Program, Kentucky Department of Fish and Wildlife Resources , 1 Sportsman’s Lane, Frankfort, Kentucky 40601 , USA