1. Horticulture College, Hunan Agricultural University, Changsha 410128, China
2. Hunan Engineering Research Center of Edible Fungi, Changsha 410128, China
3. Key Laboratory for Vegetable Biology of Hunan Province, Changsha 410128, China
4. Engineering Research Center for Horticultural Crop Germplasm Creation and New Variety Breeding, Ministry of Education, Changsha 410128, China