1. Institute of Marine Sciences, University of California, Santa Cruz, 115 McAllister Way, Santa Cruz, CA 95060, USA
2. US Geological Survey, Pacific Coastal and Marine Science Center, 2885 Mission Street, Santa Cruz, CA 95060, USA
3. School of BioSciences, Biosciences 4, The University of Melbourne, Royal Parade, Parkville, VIC 3052, Australia