1. Animal Science Department, Biomedical Primate Research Centre, Lange Kleiweg 161, 2288 GJ Rijswijk, The Netherlands
2. Department Population Health Sciences, Animals in Science & Society, Faculty of Veterinary Medicine, Utrecht University, 3584 CM Utrecht, The Netherlands
3. Department of Virology, Biomedical Primate Research Centre, Lange Kleiweg 161, 2288 GJ Rijswijk, The Netherlands