1. Department of Agroecology, Aarhus University, Blichers Allé 20, 8830 Tjele, Denmark
2. Sino-Danish College (SDC), University of Chinese Academy of Sciences, Eastern Yanqihu Campus, 380 Huaibeizhuang, Huairou, Beijing 101400, China
3. Center for Agricultural Resources Research, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Shijiazhuang 050021, China