1. Department of Plant Biotechnology, Korea University, Seoul 02841, Republic of Korea
2. Department of Horticulture, Chonnam National University, Gwangju 61186, Republic of Korea
3. Division of Biotechnology, Korea University, Seoul 02841, Republic of Korea
4. Carl R. Woese Institute for Genomic Biology, University of Illinois at Urbana-Champaign, Urbana, IL 61801, USA