1. School of Public Health, Lanzhou University, Lanzhou 730000, China
2. MOE Key Laboratory of Western China’s Environmental System, College of Earth and Environmental Sciences, Lanzhou University, Lanzhou 730000, China
3. Gansu Provincial Institute of Cultural Relics and Archaeology, Lanzhou 730000, China
4. Shenzhen Branch, Guangdong Laboratory of Lingnan Modern Agriculture, Key Laboratory of Synthetic Biology, Ministry of Agriculture and Rural Affairs, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences, Shenzhen 518120, China
5. Group of Alpine Paleoecology and Human Adaptation (ALPHA), State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China
6. Department of Genetics, University of Cambridge, Cambridge CB2 3EH, UK
7. Centre for Ancient Environmental Genomics, Globe Institute, University of Copenhagen, 1350 Copenhagen, Denmark