1. Key Laboratory of Virology and Biosafety, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan 430200, China
2. University of Chinese Academy of Sciences, Beijing 101408, China
3. Department of Microbiology and Immunology, University of Texas Medical Branch, Galveston, TX 77551, USA
4. Hubei Jiangxia Laboratory, Wuhan 430207, China