1. College of Fisheries, Hubei Hongshan Laboratory, Huazhong Agricultural University, Wuhan 430070, China
2. Engineering Research Center of Green Development for Conventional Aquatic Biological Industry in the Yangtze River Economic Belt, Ministry of Education, Wuhan 430070, China
3. Hubei Provincial Engineering Laboratory for Pond Aquaculture, Wuhan 430070, China
4. College of Life Sciences and Technology, Tarim University, Alar 843300, China