1. Enzymatic Engineering and Microbiology Laboratory, Algae Biotechnology Team, Biological Engineering Department, National School of Engineers of Sfax, University of Sfax, Sfax 3038, Tunisia
2. Université Clermont Auvergne, Clermont Auvergne INP, CNRS, Institut Pascal, F-63000 Clermont-Ferrand, France
3. INRAE, Animal Research Unit and Functionalities of Animal Products (UR AFPA), University of Lorraine, USC 340, F-54000 Nancy, France