1. School of Environmental Sciences, University of East Anglia, Norwich NR4 7TJ, UK
2. Department of Marine Environment and Engineering, National Sun Yat-Sen University, Kaohsiung 80424, Taiwan
3. Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria
4. Institute of Zoology, University of Natural Resources and Life Sciences, Gregor-Mendel-Straße 33, 1180 Vienna, Austria