1. Department of Marine Environment and Engineering, National Sun Yat-sen University, Kaohsiung 80424, Taiwan
2. School of Environmental Sciences, University of East Anglia, Norwich NR4 7TJ, UK
3. Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria
4. Department of Integrative Biology and Biodiversity Research, Institute of Zoology, University of Natural Resources and Life Sciences, Gregor-Mendel-Straße 33, 1180 Vienna, Austria