1. College of Agricultural and Environmental Sciences, Makerere University (MAK), Kampala P.O. Box 7062, Uganda
2. National Crops Resources Research Institute (NaCRRI), Kampala P.O. Box 7084, Uganda
3. College of Natural Sciences, Makerere University (MAK), Kampala P.O. Box 7062, Uganda
4. International Institute for Tropical Agriculture (IITA), Ibadan 200113, Nigeria