1. Key Laboratory of Southwest China Wildlife Resources Conservation, Ministry of Education, China West Normal University, Nanchong 637009, China
2. College of Life Sciences, China West Normal University, Nanchong 637009, China
3. Institut de Systématique, Evolution, Biodiversité, ISYEB-UMR 7205 MNHN-CNRS-Sorbonne Université-EPHE-University Antilles, Muséum National d’Histoire Naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France