1. Key Laboratory of Southwest China Wildlife Resources Conservation, Ministry of Education, China West Normal University, Nanchong 637009, China
2. College of Life Sciences, China West Normal University, Nanchong 637009, China
3. Key Laboratory of Economic Crops Genetic Improvement and Integrated Utilization, Hunan University of Science and Technology, Xiangtan 411201, China
4. Institut de Systématique, Evolution, Biodiversité, ISYEB-UMR 7205 MNHN-CNRS-Sorbonne, Université-EPHE-University Antilles, Muséum National d’Histoire Naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France