1. Department of Biology, Algoma University, Sault Ste. Marie, ON P6A2G4, Canada
2. Key Laboratory for Agro-Ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha 410125, China
3. Huanjiang Observation and Research Station for Karst Ecosystem, Chinese Academy of Sciences, Huanjiang 547100, China