1. College of Life Science, Northeast Agricultural University, Harbin 150030, China
2. College of Horticulture & Landscape Architecture, Northeast Agricultural University, Harbin 150030, China
3. Horticulture Branch, Heilongjiang Academy of Agricultural Sciences, Harbin 150040, China
4. Key Laboratory of Biology and Genetic Improvement of Horticultural Crops (Northeast Region), Ministry of Agriculture and Rural Affairs, Harbin 150030, China
5. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, China