1. College of Marine Sciences, Shanghai Ocean University, Shanghai 201306, China
2. National Engineering Research Center for Oceanic Fisheries, Shanghai 201306, China
3. Key Laboratory of Oceanic Fisheries Exploration, Ministry of Agriculture and Rural Affairs, Shanghai 201306, China
4. The Key Laboratory of Sustainable Exploitation of Oceanic Fisheries Resources, Shanghai Ocean University, Ministry of Education, Shanghai 201306, China
5. Center for Polar Research, Shanghai Ocean University, Shanghai 201306, China
6. Department of Marine Biosciences, Tokyo University of Marine Science and Technology, Minato, Tokyo 108-8477, Japan