1. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. CAS Research Center for Ecology and Environment of Central Asia, Urumqi 830011, China
4. Department of Geography, Ghent University, 9000 Ghent, Belgium