1. Key Laboratory of National Forestry and Grassland Administration on Conservation and Utilization of Warm Temperate Zone Forest and Grass Germplasm Resources, Shandong Provincial Center of Forest and Grass Germplasm Resources, Jinan 250102, China
2. College of Forestry, Shandong Agricultural University, Tai’an 271018, China
3. State-Owned Yishan Forest Farm in Yishui County, Linyi 276400, China