1. College of Fisheries and Life Science, Shanghai Ocean University, Shanghai 201306, China
2. Engineering Technology Research Center of Marine Ranching, Shanghai Ocean University, Shanghai 201306, China
3. Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, China
4. Key Laboratory of Freshwater Fisheries and Germplasm Resources Utilization, Ministry of Agriculture and Rural Affairs, Freshwater Fisheries Research Center, Chinese Academy of Fishery Sciences, Wuxi 214081, China