1. Applied Biology and Biotechnology Research Team B.P. 300, Multidisciplinary Faculty of Nador, Mohammed Premier University, Selouane 62700, Morocco
2. Conservation and Valorisation of Natural Resources Laboratory, Faculty of Sciences Dhar El Mehraz, Biotechnology, Sidi Mohamed Ben Abdellah University, Fez. B.P. 1796, Fès-Atlas 30003, Morocco
3. UMR CNRS 6553 Ecosystems, Biodiversity, Evolution (ECOBIO), University of Rennes, F-35000 Rennes, France