1. Key Laboratory of Southwest China Wildlife Resources Conservation (Ministry of Education), China West Normal University, Nanchong 637009, China
2. Daxiangling Provincial Nature Reserve, Ya’an 625200, China
3. School of Ecology and Nature Conservation, Beijing Forestry University, Beijing 100083, China
4. Key Laboratory of Artificial Propagation and Utilization in Anurans of Nanchong City, China West Normal University, Nanchong 637009, China
5. College of Panda, China West Normal University, Nanchong 637009, China