1. Faculty of Forestry and Environmental Management, University of New Brunswick, Fredericton, NB E3B 5A3, Canada
2. Faculty of Natural Resource Management, Lakehead University, Thunder Bay, ON P7B 5E1, Canada
3. Department of Biology, University of New Brunswick, Fredericton, NB E3B 5A3, Canada
4. Canadian Wildlife Service, Environment Canada, P.O. Box 6227, Sackville, NB E4L 4N1, Canada