1. Ethiopian Biodiversity Institute, Addis Ababa P.O. Box 30726, Ethiopia
2. Department of Range and Forest Resources, Botswana University of Agriculture and Natural Resources, Private Bag 0027, Gaborone P.O. Box 47204, Botswana
3. National Herbarium of Ethiopia, College of Natural and Computational Sciences, Addis Ababa University, Addis Ababa P.O. Box 3434, Ethiopia