1. Ph.D. Candidate, Dept. of Natural Resources and Environmental Management, Univ. of Hawai‘i at Mānoa, 1910 East-West Rd., Sherman Lab 101, Honolulu, Hawai‘i 96822 (corresponding author).
2. Professor, Dept. of Natural Resources and Environmental Management, Univ. of Hawai‘i at Mānoa, 1910 East-West Rd., Sherman Lab 101, Honolulu, Hawai‘i 96822.